When Morgan hybridised yellow-bodied, white-eyed females to brown-bodied, red-eyed males and intercrossed their F1 progeny, the F2 ratio deviated very significantly from the 9:3:3:1 ratio. this can be attributed to the fact that:

(1) The genes are located on X and Y chromosomes

(2) Fruit fly has abnormal chromosomes

(3) The genes are located on the X chromosome

(4) the genes exhibit incomplete dominance

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