Variations during mutations of meiotic recombinations are 

1. random and directionless
2. random and directional 
3. random and small  
4. random, small and directional

(a) Variations during mutaions of meiotic recombinations are random and directionless. Hugo de Vries based on his work on evening primrose stated that it is mutation which causes sudden appearence of variations that results in speciation.

He stated that mutations are sudden, heritable and peristant in successive generation. 

he contracdicted Darewinian variations that are small and directional.