When Morgan hybridised yellow-bodied, white-eyed females to brown-bodied, red-eyed males and intercrossed their F1 progeny, the F2 ratio deviated very significantly from the 9:3:3:1 ratio. This can be attributed to the fact that:
| 1. | The genes are located on the X and Y chromosomes. |
| 2. | Fruit fly has abnormal chromosomes. |
| 3. | The genes are located on the X chromosome. |
| 4. | The genes exhibit incomplete dominance. |